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Todoric J., Antonucci L., and Karin M., “Targeting Inflammation in Cancer Prevention and Therapy,” Cancer Prevention Research 9 (2016): 895–905, 10.1158/1940-6207.CAPR-16-0209

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Even though Tyr9 is not a base causing the unusual acidity of the thiol, it contributes to the mechanism as indicated by the low isomerase activity of the Tyr9Phe mutant, being only 2% of the wildtype activity (Pettersson and Mannervik, 2001)